Updated checklist of the genus Carabus
(includes all Carabus-species and subspecies described before 2004)
Ó Boleslav Brezina, 2003
genus: CARABUS Linné, 1758
(Digitulati)
subgenus: Carabus species: 7
Lichnocarabus 2
Eucarabus 6
Parhomopterus 11
Archaeocarabus 40
Isiocarabus 8
Ohomopterus 14
(Lepidospinulati)
Limnocarabus 1
Euleptocarabus 1
(Lipastrimorphi)
Morphocarabus 22
Trachycarabus 9
Mimocarabus 5
Cryptocarabus 10
Ophiocarabus 14
Cyclocarabus 10
Lipaster 1
(Archicarabomorphi)
Archicarabus 9
Ischnocarabus 2
Gnathocarabus 1
Acrocarabus 2
(Spinulati)
Apotomopterus 132
(Crenolimbi)
Hemicarabus 4
Homoeocarabus 1
(Latitarsi)
Neocarabus 1
Diocarabus 12
Tomocarabus 7
Stephanoarabus 1
Scambocarabus 6
Semnocarabus 8
Ulocarabus 2
Pachystus 5
Eurycarabus 2
Nesaeocarabus 2
Rhipocarabus 1
Tmesicarabus 1
Tachypus 3
Mesocarabus 4
Tanaocarabus 3
Pachycarabus 5
Cavazzutiocarabus 1
Orinocarabus 7
Oreocarabus 7
Cytilocarabus 2
Piocarabus 1
Piocarabus 2
Qinlingocarabus 5
Hypsocarabus 7
Rhigocarabus 34
Aulonocarabus 12
Leptocarabus 11
Meganebrius 22
(Arciferi)
Platycarabus 5
Chaetocarabus 2
Heterocarabus 1
(Procrustimorphi)
Hygrocarabus 1
Ctenocarabus 2
Cechenochilus 2
Procechenochilus 3
Microtribax 3
Tribax 17
Neoplectes 2
Archiplectes 18
Microplectes 2
Sphodristocarabus 18
Oxycarabus 1
Procrustes 10
Lamprostus 12
Cathoplius 1
Macrothorax 3
Relictocarabus 1
Apoplesius 4
Iniopachys 2
Chrysotribax 2
Chrysocarabus 5
Procerus 3
Megodontus 18/1820
Imaibius 21
Deroplectes 5
Axinocarabus 1
Goniocarabus 7
Eotribax 10
Alipaster 7
Cratocechenodes 1
Cechenotribax 1
Cratocarabus 5
Cratocephalus 6
Cratocechenus 4
Cratophyrtus 6
Pseudotribax 2
Pantophyrtus 5
Leptoplesius 8
Pseudocranion 25
Shunichiocarabus 1
Cupreocarabus 10
Neoplesius 30
Eocechenus 9
Calocarabus 12
Cryptocechenus 2
Aristocarabus 1
Pagocarabus 1
Megodontoides 4
Pseudocoptolabrus 6
Imaibiodes 1
Acathaicus 1
Cathaicus 1
Eupachys 2
Fulgenticarabus 1
Teratocarabus 1
Cychrostomus 2
Cephalornis 1
Eccoptolabrus 1
Lasiocoptolabrus 1
Coptolabrus 14
Coptolabrodes 1
Shenocoptolabrus 1
Acoptolabrus 7
Damaster 1
subgenera: 118
species: 823/584/141852
Carabus Linné, 1758
subgenera: 118
species: 823/584/141852
(Digitulati)
carabus Linné, 1758; type species Carabus granulatus Linné
species: 7
1 arcensisarvensis Herbst, 1784; cosemend. for arcensis Herbst
ssp. arcensisarvensis Herbst, 1784; cosemend. for arcensis Herbst
ssp. conciliator Fischer, 1822
ssp. faldermanni Dejean, 1829
ssp. hokkaidoensis Lapouge, 1924
ssp. baschkiricus Breuning, 1936
ssp. sylvaticus Dejean, 1826
ssp. carpathus Born, 1902; ?non priority
ssp. venetianus Bernau, 1914
2 deyrollei Gory, 1839
3 granulatus Linné, 1758
ssp. granulatus Linné, 1758
ssp. duarius Fischer, 1844; nom.pro elongatus Fischer
ssp. interstitialis Duftschmid, 1812
ssp. corticalis Motschulsky, 1846, nec Fabricius, 1801; lsu
ssp. leander Kraatz, 1878
ssp. yezoensis Bates, 1883
ssp. telluris Bates, 1883
4 menetriesi Hummel, 1827
ssp. menetriesi Hummel, 1827
ssp. pacholei Sokolar, 1911
5 sculpturatus Ménétriés, 1832
6 stscheglowi Mannerheim, 1827
7 vanvolxemi Putzeys, 1875
Lichnocarabus Reitter, 1896; type species Carabus vinctus Weber
species: 2
1 limbatus Say, 1825, nec Fabricius, 1776; lsu
2 vinctus Weber, 1801
Eucarabus Géhin, 1885; type species Carabus ullrichi Germar
species: 6
1 catenulatus Scopoli, 1763
ssp. catenulatus Scopoli, 1763
ssp. fontanellae Reitter, 1896
2 cumanus Fischer, 1823
3 italicus Dejean, 1826
ssp. italicus Dejean, 1826
ssp. rostagnoi Luigioni, 1904
4 obsoletus Sturm, 1815, nec Olivier, 1795; lsu
ssp. obsoletus Sturm, 1815, nec Olivier, 1795; lsu
ssp. carpathicus Palliardi, 1825
ssp. nagyagensis Seidlitz, 1888; nom.pro procerus Bielz
5 parreyssi Palliardi, 1825; ?iss of paraysi Palliardi
ssp. parreyssi Palliardi, 1825; ?iss of paraysi Palliardi
ssp. plassensis Born, 1907
6 ullrichi Germar, 1824; ?iss of ulrichii Germar
ssp. ullrichi Germar, 1824; ?iss of ulrichii Germar
ssp. rhilensis Kraatz, 1876
ssp. fastuosus Palliardi, 1825
ssp. arrogans Schaum, 1858
Parhomopterus Lapouge, 1931; type species Carabus sternbergi Roeschke
species: 11
1 angustus Roeschke, 1898
2 billbergi Mannerheim, 1827
ssp. billbergi Mannerheim, 1827
ssp. coreicola Deuve, 1989
ssp. lupisylvaticus Imura, 1993
ssp. jilinicola Deuve & Li, 1999
3 cartereti Deuve, 1982
ssp. cartereti Deuve, 1982
ssp. ronkayi Deuve, 1992
ssp. omisanensis Lassalle, 1999
4 manifestus Kraatz, 1881
ssp. manifestus Kraatz, 1881
ssp. guanmenshanus Imura, 1991
ssp. pyonganicola Deuve & Li, 1998
5 mianyangensis Deuve & Li, 1998
6 namhaedoensis Kwon & Lee, 1984
7 nitididorsus Ishikawa & Kim, 1983
ssp. nitididorsus Ishikawa & Kim, 1983
ssp. paegunsanus Imura, 1997
8 pawlowskianus Deuve, 1989
ssp. pawlowskianus Deuve, 1989
ssp. peregrinus Cavazzuti & Ratti, 1998
9 sternbergi Roeschke, 1898
ssp. sternbergi Roeschke, 1898
ssp. lauschanensis Born, 1913
ssp. kojedoensis Kwon & Lee, 1984
ssp. tongdosanus Kwon & Lee, 1984
10 szeli Deuve, 1994
11 xiuyanensis Deuve & Li, 1998
Archaeocarabus Semenov, 1898; type species Carabus relictus Semenov
species: 40
1 agamemnon Breuning, 1943
2 anhuinus Imura, 1996
3 baimanorum Deuve, 1999
4 battoniensis Deuve, 1991
5 bornianus Hauser, 1922
6 camilloi Cavazzuti & Ratti, 1998
7 claricollis Cavazzuti & Ratti, 1998
8 daiyunshan Kleinfeld, 1998
9 dongchuanicus Deuve, 1994
ssp. dongchuanicus Deuve, 1994
ssp. mianshanicus Cavazzuti, 1997
ssp. luojicus Cavazzuti, 1998
ssp. meiguensis Imura, 1998
10 dreuxioides Deuve, 1998
11 emanuelei Imura, 1997
12 fontellus Deuve, 1997
ssp. fontellus Deuve, 1997
ssp. semifuscus Cavazzuti & Ratti, 1999
ssp. attrepidans Deuve, 2000
ssp. pseudoemeicola Deuve, 2001
13 gressittianus Mandl, 1975
ssp. gressittianus Mandl, 1975
ssp. fuscomarginatus Kleinfeld, 1997
ssp. shanminicus Deuve, 1999
ssp. bambousicola Deuve & Tian, 1999
ssp. daanensis Deuve, 1999
14 koidei Imura, 1996
15 kucerai Deuve, 1997
16 kweitshauensis Mandl, 1975
17 latens Cavazzuti & Ratti, 1998
18 latipennis Breuning, 1932
ssp. latipennis Breuning, 1932
ssp. shansutiaensis Lassalle & Remond, 1998
19 ludingensis Deuve & Vigna Taglianti, 1992
ssp. ludingensis Deuve & Vigna Taglianti, 1992
ssp. gonggaiensis Deuve, 2002
20 maowenensis Deuve, 2001
21 mianningensis Kleinfeld, 2000
22 morphocaraboides Deuve, 1989
ssp. morphocaraboides Deuve, 1989
ssp. dongchuan Cavazzuti, 2001
ssp. pigmentatus Cavazzuti, 1998
ssp. yao Imura, 1999
ssp. zhaojueanus Imura, 1999
23 nanosomus Hauser, 1931
ssp. nanosomus Hauser, 1931
ssp. cordithoracicus Deuve, 1989
ssp. maotaiophilus Deuve, 1996
ssp. wufengensis Kleinfeld, 1998
24 nestor Breuning, 1934
ssp. nestor Breuning, 1934
ssp. xianfengicus Deuve & Li, 2000
=xianfeng Kleinfeld, 2000 SW-Hubei:Xianfeng, 29:42N/108:58E:Laoyan
ssp. emeicola Deuve & Imura, 1992
ssp. barbulatus Cavazzuti & Ratti, 1999
25 oblongior Deuve, 1992
ssp. oblongior Deuve, 1992
ssp. yunlingicus Deuve, 1994
26 pani Deuve & Tian, 2001
27 paris Breuning, 1932
28 pseudolatipennis Deuve, 1991
ssp. pseudolatipennis Deuve, 1991
ssp. nanpingensis Deuve, 1992
ssp. wenxianensis Deuve, 1992
ssp. bashanensis Imura, 1995
ssp. shennongjianus Imura, 1996
ssp. qinlingicus Deuve, 1998
ssp. huayangzhen Cavazzuti, 1999
ssp. lantianensis Cavazzuti, 1999
ssp. yingpanjienus Cavazzuti, 1999
ssp. yangxianensis Deuve, 1999
ssp. gracilescens Deuve, 1999
29 relictus Semenov, 1898
30 shamaevi Imura, 1996
31 tianbaoshan Kleinfeld, 1998
32 tieguanzi Imura, 1990
33 vigil Semenov, 1898; nom.pro striatus Semenov
ssp. vigil Semenov, 1898; nom.pro striatus Semenov
ssp. pseudoparis Deuve, 1992
ssp. dabashanus Imura, 1995
ssp. nigrominimus Deuve & Mourzine, 1997
ssp. pseudotosanus Deuve & Mourzine, 1997
ssp. pyrochromus Deuve, 1997
ssp. quercinus Cavazzuti, 1999
ssp. cordulatus Cavazzuti, 1999
ssp. guangwushanus Imura & Su, 2000
ssp. xunyangbaensis Deuve, 2000
ssp. ohshimaorum Deuve, 1991
ssp. giacomazzoi Deuve, 1992
34 vigilax Bates, 1890
ssp. vigilax Bates, 1890
ssp. wassulandensis Deuve, 2000
35 wuangxianus Kleinfeld, 1998
36 wushanling Kleinfeld, 1998
37 wutaishanicus Müller, 2000
38 xingshanensis Imura, 1996
39 xupuensis Kleinfeld, 1998
40 yunnanus Fairmaire, 1886
ssp. yunnanus Fairmaire, 1886
ssp. guerryanus Breuning, 1961
ssp. pseudoyunnanus Deuve, 1994
ssp. yanyuanicus Cavazzuti, 1996
ssp. hei Cavazzuti, 1996
Isiocarabus Reitter, 1896; type species Carabus fiduciarius Thomson
species: 8
1 fiduciarius Thomson, 1856
ssp. fiduciarius Thomson, 1856
ssp. kirinicus Csiki, 1927; nom.pro auricollis Born
ssp. saishutoicus Csiki, 1927; nom.pro insularis Lapouge
ssp. tim Kleinfeld, 1999
2 hienfoungi Thomson, 1857
ssp. hienfoungi Thomson, 1857
ssp. chinensis Breuning, 1927
3 hunanicola Deuve & Yu, 1992
ssp. hunanicola Deuve & Yu, 1992
ssp. dreuxi Deuve, 1994
ssp. dyseimatus Cavazzuti, 1995
ssp. jinxiuensis Imura, 1995
ssp. yangmingensis Deuve, 1995
ssp. liangshanicus Kleinfeld, 1998
ssp. loccaianicolor Deuve & Tian, 1999
ssp. guandongicola Deuve & Tian, 2000
ssp. yulingensis Kleinfeld, 2002
4 kiukiangensis Bates, 1888
ssp. kiukiangensis Bates, 1888
ssp. chusanensis Nicolas, 1924
ssp. alzonai Deuve, 1990
ssp. ronganensis Deuve, 1998
ssp. pseudoronganensis Deuve & Tian, 2000
ssp. orphniopterus Hauser, 1920; nom.pro castanopterus Hauser
ssp. fulvoligatus Cavazzuti & Ratti, 1998
5 miaorum Lassalle & Prunier, 1993
6 pustululatus Deuve, 1993
7 strandiellus Breuning, 1934
ssp. strandiellus Breuning, 1934
ssp. heterodynamusDeuve, 1991
ssp. tianzhuicus Imura, 1996
ssp. microgranum Cavazzuti & Ratti, 1999
ssp. guangchangensis Deuve, 2001
8 dargei Deuve, 1987
Ohomopterus Reitter, 1896; type species Carabus dehanii Chaudoir
species: 14
1 albrechti Morawitz, 1862
ssp. albrechti Morawitz, 1862
ssp. freyi van Emden, 1932
ssp. esakianus Nakane, 1961
ssp. okumurai (Ishikawa, 1966)
ssp. tohokuensis Ishikawa, 1984
ssp. awashimae Ishikawa & Takami, 1996
ssp. echigo Ishikawa & Takami, 1996
ssp. hidakanus Ishikawa & Takami, 1996
ssp. itoi Ishikawa & Takami, 1996
2 arrowianus Breuning, 1934
ssp. arrowianus Breuning, 1934
ssp. nakamurai (Ishikawa, 1966)
ssp. komiyai (Ishikawa, 1966)
ssp. murakii Ishikawa & Kubota, 1984
ssp. hidaosa Ishikawa & Kubota, 1995
ssp. minoensis Ishikawa & Kubota, 1995
3 daisen (Nakane, 1953)
ssp. daisen (Nakane, 1953)
ssp. okianus (Nakane, 1961)
4 dehanii Chaudoir, 1848
ssp. dehanii Chaudoir, 1848
ssp. ishidai Imura & Mizusawa, 1995
ssp. katsumai Imura & Mizusawa, 1995
ssp. koshikicola Imura & Mizusawa, 1995
ssp. kumaso Imura & Mizusawa, 1995
ssp. tosanus (Nakane, Iga & Ueno, 1953)
ssp. ishizuchianus (Nakane, 1960)
ssp. kawanoi (Kamiyoshi & Mizoguchi, 1960)
ssp. botchan Imura & Mizusawa, 1995
5 esakii Csiki, 1927; nom.pro lewisi Géhin & Haury
6 insulicola Chaudoir, 1869
ssp. insulicola Chaudoir, 1869
ssp. awashimaensis Ishikawa & Ujiie, 2000
ssp. kantoensis Ishikawa & Ujiie, 2000
ssp. kita Ishikawa & Ujiie, 2000
ssp. okutonensis Ishikawa & Ujiie, 2000
ssp. shinano Ishikawa & Ujiie, 2000
ssp. kiso Ishikawa & Ujiie, 2000
ssp. komaganesis Ishikawa & Ujiie, 2000
ssp. sado Ishikawa & Ujiie, 2000
7 iwawakianus (Nakane, 1953)
ssp. iwawakianus (Nakane, 1953)
ssp. kiiensis (Nakane & Iga, 1953)
ssp. muro Ishikawa & Kubota, 1996
ssp. narukawai Ishikawa & Kubota, 1996
ssp. shima Ishikawa & Kubota, 1996
8 japonicus Motschulsky, 1857
ssp. japonicus Motschulsky, 1857
ssp. tsushimae Breuning, 1932; emend.pro tsuchimae Breuning, 1932
ssp. chugokuensis (Nakane, 1961)
ssp. ikiensis (Nakane, 1968)
ssp. chotaroi Imura, Dejima & Mizusawa, 1993
ssp. hiradonis Imura, Dejima & Mizusawa, 1993
ssp. onodai Imura, Dejima & Mizusawa, 1993
ssp. okinoshimanus Imura, Dejima & Mizusawa, 1993
ssp. awajiensis Imura, Dejima & Mizusawa, 1993
ssp. seizaburoi Imura, Dejima & Mizusawa, 1993
ssp. nozakicola Imura & Mizusawa, 1994
ssp. wakamatsuensis Imura & Mizusawa, 1994
ssp. mochizukii Imura & Mizusawa, 1994
ssp. umekii Imura & Mizusawa, 1999
ssp. yoshiyukii Imura & Mizusawa, 1999
9 kimurai (Ishikawa, 1966)
10 lewisianus Breuning, 1932
ssp. lewisianus Breuning, 1932
ssp. awakazusanus Ishikawa, 1981
11 maiyasanus Bates, 1873
ssp. maiyasanus Bates, 1873
ssp. hokurikuensis Ishikawa & Kubota, 1995
ssp. yoroensis Ishikawa & Kubota, 1995
ssp. ohkawai (Nakane, 1968)
ssp. shigaraki (Hiura & Katsura, 1971)
ssp. takiharensis Katsura & Tominaga, 1978
ssp. suzukanus Ishikawa & Kubota, 1995
12 uenoi (Ishikawa, 1960)
13 yaconinus Bates, 1873
ssp. yaconinus Bates, 1873
ssp. blairi Breuning, 1934
ssp. cupidicornis Ishikawa & Kubota, 1994
ssp. maetai Ishikawa & Kubota, 1994
ssp. oki Ishikawa & Kubota, 1994
ssp. seto Ishikawa & Kubota, 1994
ssp. sotai Ishikawa & Kubota, 1994
ssp. yamaokai Ishikawa & Kubota, 1994
14 yamato (Nakane, 1953)
(Lepidospinulati)
Limnocarabus Géhin, 1876; type species Carabus clathratus Linné
species: 1
1 clathratus Linné, 1761
ssp. clathratus Linné, 1761
ssp. stygius Ganglbauer, 1890
ssp. auraniensis Müller, 1902
ssp. maacki Morawitz, 1862
Euleptocarabus Nakane, 1956; type species Carabus porrecticollis Bates
species: 1
1 porrecticollis Bates, 1883
ssp. porrecticollis Bates, 1883
ssp. kansaiensis Nakane, 1961
(Lipastrimorphi)
Morphocarabus Géhin, 1885; type species Carabus monilis Fabricius
species: 22
1 aeruginosus Fischer, 1822
ssp. aeruginosus Fischer, 1822
ssp. hermani Mannerheim, 1827
ssp. salechardensis Obydov, 1999
ssp. eschscholtzi Mannerheim, 1827
ssp. tulatensis O.Berlov & Shilenkov, 1996
ssp. zyrjanovskianus Shilenkov & O.Berlov, 1996
ssp. verzhutzkii O.Berlov & Shilenkov, 1996
ssp. kolyvanus Shilenkov & O.Berlov, 1996
2 chaudoiri Gebler, 1847
ssp. chaudoiri Gebler, 1847
ssp. mongolorum Csiki, 1927; nom.pro neglectus Lapouge, 1913
ssp. czadanicus Obydov, 1997
ssp. marusiki Obydov, 1997
ssp. zaikai Obydov, 1999
ssp. beladici Obydov, 2001
ssp. shilenkovi O.Berlov, 1989
3 czernyscheviellus Obydov, 2002
4 excellens Fabricius, 1798
5 gebleri Fischer, 1817
ssp. gebleri Fischer, 1817
ssp. ultimus Obydov, 1999
6 henningi Fischer, 1817
ssp. henningi Fischer, 1817
ssp. oviformis Beheim & Breuning, 1943
ssp. hamarensis Shilenkov, 1996
ssp. melyachi Shilenkov & Menstshikov, 1996
ssp. saldaitisi Obydov, 1997
ssp. peczoricus Obydov, 2000
7 hummeli Fischer, 1823
ssp. hummeli Fischer, 1823
ssp. middendorfi Ménétriés, 1851
ssp. stolidus Lapouge, 1924
ssp. jurgae Obydov, 1995
ssp. nevelskii Shilenkov, 1996
ssp. smaragdulus Kraatz, 1878, nec Fabricius; 1798; lsu
ssp. tristiculus Kraatz, 1879
ssp. pusongensis Imura, 1993
ssp. suensoni Mandl, 1979
8 karpinskii Kryzhanovskij & Matveev, 1993
9 kozhantschikovi Lutshnik, 1924
10 laojunensis Kleinfeld, 1999
11 mestscherjakovi Lutshnik, 1924
ssp. mestscherjakovi Lutshnik, 1924
ssp. pseudoodoratus Brinev, 2002
ssp. zherichini Shilenkov, 1990
ssp. shaman Shilenkov, 1996
ssp. telezkianus Shilenkov, 1996
12 michailovi Kabak, 1992
13 monilis Fabricius, 1792
14 odoratus Motschulsky, 1844; ?non priority
ssp. odoratus Motschulsky, 1844
ssp. viridilimbatus Motschulsky, 1845
ssp. irkoutskensis Lapouge, 1915
ssp. dohrni Gebler, 1847; ?non priority
ssp. baeri Ménétriés, 1851
ssp. chaffanjoni Lesne, 1897
ssp. melleus Lapouge, 1909; ?un
ssp. septentrionalis Breuning, 1932
ssp. kamtschatensis Breuning, 1942
ssp. antropovi Shilenkov, 1996
ssp. bargusinicus Shilenkov, 1996
ssp. dabanensis Shilenkov, 1996
ssp. krugeri Obydov, 1999
ssp. magadanicus Obydov, 1999
ssp. martjanovianus Obydov, 1999
ssp. taskylensis Obydov, 2001
15 regalis Fischer, 1822
ssp. regalis Fischer, 1822
ssp. jenissoni Dejean, 1831
16 rothi Dejean, 1829
ssp. rothi Dejean, 1829
ssp. comptus Dejean, 1831
ssp. hampei Küster, 1846
ssp. incompsus Kraatz, 1880
ssp. alutensis Savulescu, 1972
17 scheidleri Panzer, 1799
ssp. scheidleri Panzer, 1799
ssp. helleri Ganglbauer, 1893; ?non priority
ssp. kollari Palliardi, 1825
ssp. praecellens Palliardi, 1825
ssp. illigeri Dejean, 1826, nec Panzer, 1806; lsu
ssp. versicolor Frivaldsky, 1835
ssp. zawadzkii Kraatz, 1854
ssp. seriatissimus Reitter, 1896
ssp. ronayi Csiki, 1906
ssp. simulator Kraatz, 1876
18 spasskianus Fischer, 1823
ssp. spasskianus Breuning, 1932
ssp. tungus Obydov, 1999
ssp. putus Motschulsky, 1846
ssp. schestopalovi Plutenko, 1995
ssp. geblerianus Obydov, 1999