HACKBERRY

Many tree species can be easily identified by their bark alone. One tree species has bark that resembles worms crawling upon it. That tree species is the Hackberry (Celtis occidentalis L.).

This tree is a member of the Order Rosales and of the Family Cannabaceae, Celtidaceae, or Ulmaceae. (Different references list different families.)

The generic name, Celtis, is a Latin name given by Roman naturalist Pliny the Elder to an unrelated Lotus tree, which may have been called “Celthis”. The specific epithet, occidentalis, refers to the Western Hemisphere. Other scientific synonyms for this species are Celtis canina Rafinesque and Celtis pumila Pursh.

The common name, Hackberry, is from the Scottish “Hagberry”, which means “Marshberry”. That is the name for their Bird Cherry (Prunus avium [L.] L.) tree that grows in their wetlands, called “hags”. Other common names for this species are American Hackberry, Bastard Elm, Beaverwood, Common Hackberry, False Elm, Hack-tree, Hoop Ash, Nettletree, Northern Hackberry, Sugarberry, and Witches’ Broom. This tree is both fast growing and moderately long-lived.

DESCRIPTION OF THE HACKBERRY

Size:

Height: Its height is 20-130 feet.

Trunk Diameter: Its trunk diameter is 10-54 inches.

Crown: Its crown is open, symmetrical, spreading, rounded, oblong, and broadly columnar. Its branches are stout, spreading, and arching.

Trunk: Its trunk is both short and straight.

Twigs: Its twigs are slender; zigzag; rounded; smooth; and greenish, gray-brown, or light red-brown with white lenticels. Its leaf scars are small, crescent-shaped or semi-oval, slightly raised, and have 1-3 bundle scars. Its pith is small, white, and is finely diaphragmed or chambered at the nodes.

Buds: Its lateral buds are ¼ inch long, gray to light brown, sessile, ovate, triangular, acute, hairy, and are appressed against the twig. They have 4-5 sets of 2-ranked scales. There are no true terminal buds.

Leaves: Its leaves are deciduous, simple, and alternate. They are arranged in 2 horizontal rows along the branchlets. Each leaf is ovate to oblong lanceolate; 2-7 inches long; 1½-2½ inches wide; has a long, narrow, and slightly curved, acuminate tip; an asymmetrical, rounded or cordated, and toothless base; 3 prominent veins; and singly coarse to sharply serrated teeth near the upper 2/3 of the margin. They are glossy; papery; dark green, smooth, or slightly rough above; and are paler, smooth, or slightly hairy below. Its petiole is slender, about ¾-1 inch long, slightly grooved, and hairy. In the fall these leaves turn yellow-green to yellow-brown.

A few animal species eat these leaves. The Butterfly larvae of the American Snout (Libytheana carineta Cramer), the Question Mark (Polygonia interrogationis Fabricius), and the Hackberry Emperor (Asterocampa celtis Boisduval & Leconte) all eat these leaves. White-tailed Deer (Odocoileus virginianus Zimmermann) also eat these leaves.

Flowers: Its male flowers are arranged in small clusters upon hanging stalks at the base of the annual shoots. Its female flowers are singly arranged at the base of the upper leaves upon the same shoots as the male flowers. These flowers are either unisexual or bisexual and are monoecious. Each flower is creamy grayish-green, about ¼ inch wide, and has a 4-5 lobed calyx, and no corolla. The male flowers have 5 stamens and the female flowers have a 2-lobed style and a single-celled ovary. These flowers are wind-pollinated. Flowering season is April to May.

Fruits: Its fruits are solitarily arranged upon 1/8-1 inch, slender stems. Each fruit is a ¼-3/4 inch long, shiny, spherical drupe with a thick, tough skin and a thin, yellow-orange, fleshy pulp. They are wrinkled when dry. They are initially greenish but become orange-red or brown-purple when ripe. The fruit has 1 hard, pale brown, rounded nutlet seed with a rough surface. These seeds are dispersed by animals and by water. Fruiting season is September to November. The fruits remain upon the tree throughout the winter.

These fruits are eaten by many species of birds, such as American Crows (Corvus brachyrhynchos Brehm), American Robins (Turdus migratorius L.), Brown Thrashers (Toxostoma rufum L.), Cedar Waxwings (Bombycilla cedrorum Vieillot), Gray Catbirds (Dumetella carolinensis L.), Mourning Doves (Zenaida macroura L.), Northern Bobwhite (Colinus virginianus L.), Northern Mockingbirds (Mimus polyglottos L.), Wild Turkeys (Meleagris gallopavo L.), and Woodpeckers (Family Picidae). The fruits are also eaten by many species of Mammals (Class Mammalia), such as Fox Squirrels (Sciurus niger L.), Flying Squirrels (Genus Glaucomys), Gray Foxes (Urocyon cinereoargenteus Schreber), Mice (Genus Mus), Raccoons (Procyon lotor L.), Striped Skunks (Mephitis mephitis Schreber), and Virginia Opossums (Didelphis virginiana Kerr).

Bark: Its young bark is smooth and is light yellow-brown or gray-brown. Its older bark is scaly and has narrow, dark gray woody and corky knobs, warts, and ridges. This bark is thin and is susceptible to fire damage.

Wood: Its sapwood is thick, light yellow, greenish gray, or green yellow. Its darker heartwood is light yellowish gray to light brown. This wood is heavy, weak, soft to moderately hard, flexible, shock-resistant, ring-porous, and coarse-grained. It has distinct annual rings. It is subject to rotting when exposed to the elements.

Roots: Its fibrous roots are both deep and widespreading. They may extend down to 20 feet. This species can tolerate droughts.

Habitat: Its habitats consist of woodlands and open places. They may be found along streams and rivers, in floodplains, and in bottomlands. They are also found in moist, alkaline-rich upland soils. They can tolerate poor soils. They are rarely found in pure stands. It is moderately shade tolerant.

Range: Its range covers much of the U.S. from southern New England and Virginia to the Great Plains and north into southeastern Canada. It is not found in the Deep South and in the northern Great Lakes.

Pests and Diseases:

Hackberries are susceptible to Eriophyid Gall Mites (Eriophyes celtis Kendall) and Powdery Mildew Fungi (Sphaerotheca phytoptophila Kellerman & Swingle) that cause “Witches Brooms”. This distorts, gnarls, and twists branches and twigs but does not kill the tree. It is more noticeable when the leaves are absent. They also make ideal nesting sites for Songbirds.

The undersides of the leaves have small, rounded fleshy Nipple Galls, caused by Hackberry Nipple Gall Psyllids (Pachypsylla celtidismamma [Riley]). The Hackberry Lace Bug (Corythucha celtides Osborn & Drake) can cause premature yellowing or browning of the leaves. Unlike other members of the Elm Family, Hackberries are not affected by the Dutch Elm Disease (Ophiostama ulmi [Buisman] and Ophiostama novo-ulmi Brasier).

Hackberries have their own defense mechanism. These trees exude a gummy substance that protects their wounds from infections and from infestations.

Uses:

Hackberry has many different uses. Both the Native Americans and the early European settlers had uses for this tree.

The fruits are both sweet and edible. The soft, white kernel inside of the hard nutlet shell is also sweet and edible. These fruits can be eaten straight or can be used to sweeten other foods. They are most palatable during the late fall and winter and after a few hard, heavy frosts. The quality and abundance of these fruits depended upon their growing conditions.

Hackberry has some medicinal uses, too. A decoction of the bark, sometimes mixed with other ingredients, can be used for treating sore throats, venereal diseases, and various female disorders.

The wood is used for fuel, fence posts, boxes, crates, pallets, barrel hoops, bows, hoe handles, sporting goods, plywood, kitchen cabinets, and cheap furniture. Much of the exposed wood upon the furniture is covered with veneer from other wood species.

The bark is fibrous. It was used for making cordage and sandals.

Hackberry has other uses, too. It is used as an ornamental shade tree because it tolerates some urban air pollution. It is also used for erosion control, shelterbelts, and windbreaks.

REFERENCES

FEASTING FREE ON WILD EDIBLES

By Bradford Angier

MICHIGAN TREES

By Burton V. Barnes and Warren H. Wagner, Jr.

AN ECLECTIC GUIDE TO TREES EAST OF THE ROCKIES

By Glen Blouin

THE ENCYCLOPEDIA OF EDIBLE PLANTS OF NORTH AMERICA

By Francois Couplan, Ph. D.

THE BOOK OF FOREST AND THICKET

By John Eastman and Amelia Hansen

TREES OF THE NORTHERN UNITED STATES AND CANADA

By John Laird Farrar

TREES OF PENNSYLVANIA AND THE NORTHEAST

By Charles Fergus and Amelia Hansen

EDIBLE WILD PLANTS OF EASTERN NORTH AMERICA

By Merritt Lyndon Fernald and Alfred Charles Kinsey

TREES OF THE EASTERN AND CENTRAL UNITED STATES AND CANADA

By William M. Harlow

EAT THE WEEDS

By Ben Charles Harris

101 TREES OF INDIANA

By Marion T. Jackson

NATIONAL WILDLIFE FEDERATION FIELD GUIDE TO TREES OF NORTH AMERICA

By Bruce Kershner, Daniel Mathews, Gil Nelson, and Richard Spellenberg

AUTUMN LEAVES

By Ronald Lanner

TREES OF THE CENTRAL HARDWOOD FORESTS OF NORTH AMERICA

By Donald J. Leopold, William C. McComb, and Robert N. Muller

NATIONAL AUDUBON SOCIETY FIELD GUIDE TO NORTH AMERICAN TREES (EASTERN REGION)

By Elbert L. Little

THE FOLKLORE OF TREES AND SHRUBS

By Laura C. Martin

NATIVE AMERICAN MEDICINAL PLANTS

By Daniel E. Moerman

A NATURAL HISTORY OF TREES OF EASTERN AND CENTRAL NORTH AMERICA

By Donald Culross Peattie

EDIBLE WILD PLANTS

By Lee Allen Peterson

EASTERN TREES

By George A. Petrides

THE URBAN TREE BOOK

By Arthur Plotnik

THE SIBLEY GUIDE TO TREES

By David Allen Sibley

OHIO TREES

By T. Davis Sydnor and William F. Cowen

THE USES OF WILD PLANTS

By Frank Tozer

WINTER BOTANY

By William Trelease

NATIVE TREES OF THE MIDWEST

By Sally S. Weeks, Harmon P. Weeks, and George R. Parker

BARK: A FIELD GUIDE TO TREES OF THE NORTHEAST

By Michael Wojtech

en.wikipedia.org/wiki/Celtis_occidentalis