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I. Phylum Cnidaria- hydra, jellyfish, and corals.

A. fresh water & mostly marine

B. benthic colonial & planktonic

C. diagnostic features

1. radiate, eumetazoans

2. tissue grade

3. diploblastic

4. predaceous

a. cnidocyte- stinging cells with nematocysts

1. cnidocils in hydrozoa & scyphozoa

2. protein toxin

b. tentacles

c. sensory organs: statocysts & ocelli

5. gastro-vascular cavity- plays an important roll as a hydrostatic skeleton

6. polymorphic (polyp & madusae forms)

7. oral/aboral surfaces

8. manubrium- bears the mouth

9. vellum- shelf of tissue extending from the margin of the bell to the manubrium; provides jet propulsion

II. General Structure & Function:

A. polymorphic: fig. 4-5

1. medusoid

2. polypoid

3. alternation of generations

B. histology:

1. epidermis- contain myoepithelial cells

2. gastrodermis- lines the gastrovascular cavity nutritive cells; flagellated; phagocytic

3. mesoglea- secreted layer between epidermis and the gastrodermis; gelatinous consistency

C. nutrition/feeding: carnivores w/ extracellular digestion & phagocytosis; zoolchlorellae or zooxanthellae

D. nervous system: "nerve net" w/ receptor cells

E. respiration/excretion: diffusion/ammonia

F. reproduction: budding; sexual reproduction in medusae; planula larvae

1. Hyda- sexual reproduction occurs in fall to produce winter eggs.

2. many are dioecious

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III. Classification:

A. Class Hydrozoa (hydroids); 2700 spp.

1. mostly colonial polyps; Obelia, fig. 4-15

2. Physalia- Portuguese Man-O-War; colonial pelagic hydrozoan; pneumatophores, fig. 4-25

3. Hydra- solitary polyps

4. Craspedacusta; found in Oklahoma; contains a symbiotic algae Chlorella

B. Class Scyphozoa (true jellyfish); 200 spp.

1. mostly marine

2. medusae dominance

3. Some dangerous species

4. Cassiopea- swims upside down in shallow intertidal pools

5. Aurelia; fig. 4-35

medusa--->gametes--->zygote--->planula--->scyphistoma--->strobilus--->ephyra

C. Class Anthozoa (anemones, corals, & sea fans); 6000+ spp.

1. all marine

2. solitary or colonial forms; polyp form only

3. gastrodermis bears nematocysts w/o opercula

4. some hermaphrodites

5. stony corals produce CaCO3 skeletons; sclerosepta

6. often contain zooxanthellae

7. octocorals: sea pansies, sea fans, whip corals, & pipe corals

a. eight pinnate tentacles

b. Gorgonians figs. 4-52 & 53

c. O. Stolonifera- organpipe coral- polyps encased in parallel calcareous tubes

8. coral reefs support diverse associations of marine organisms

9. structurally heterogeneous habitat

IV. Ctenophora- comb jellies ~50 spp.

A. poorly known group

B. delicate

C. Comb rows; figs. 4-66 & 67